The daughter chromatids are optionally coiled round one another at the metaphase of mitosis. The direction of coiling is usually consistent in each arm of all chromosomes in Fritillaria and constant for the identifiable M chromosomes. In Nomocharis however the direction is sometimes inconsistent in particular arms.2. Chromatid coiling at metaphase seems to be developed during prophase chiefly as a result of a strain imposed an the chromatids by spiralisation but subject also to other conditions that have not yet been ascertained.
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Cyril Dean Darlington (1936) studied this question.
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