The tribe Osirini, its scope, classification, and revisions of the genera Parepeolus and Osirinus (Hymenoptera, Apoidea, Anthophoridae)

The limits of the hitherto monotypic tribe Osirini are expanded to include Epeoloides, Parepeolus and three new genera: Ecclitodes, Osirinus and Protosiris.It is suggested that the tribe does not belong in the subfamily Nomadinae, but represents a group of parasites independently derived from pollen-collecting ancestors.A cladistic analysis, a key and descriptions are presented for the genera.For Parepeolus and Osirinus keys to the species, synonymies and descriptions are provided; four new species are described.