R. E. Green, G. J. M. Hirons, A. R. Johnson, The Origin of Long-Term Cohort Differences in the Distribution of Greater Flamingos Phoenicopterus ruber roseus in Winter, Journal of Animal Ecology, Vol. 58, No. 2 (Jun., 1989), pp. 543-555
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Green et al. (1989) studied this question.
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