Myong Gi Chung, J. L. Hamrick, Samuel B. Jones, Jr., Gregory S. Derda, Isozyme Variation Within and Among Populations of Hosta (Liliaceae) in Korea, Systematic Botany, Vol. 16, No. 4 (Oct. - Dec., 1991), pp. 667-684
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Chung et al. (1991) studied this question.