J. D. Goss-Custard, The Ecology of the Wash. III. Density-Related Behaviour and the Possible Effects of a Loss of Feeding Grounds on Wading Birds (Charadrii), Journal of Applied Ecology, Vol. 14, No. 3 (Dec., 1977), pp. 721-739
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JOHN D. GOSS‐CUSTARD (1977) studied this question.
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