E. I. Hazard, T. G. Andreadis, D. J. Joslyn, E. A. Ellis, Meiosis and Its Implications in the Life Cycles of Amblyospora and Parathelohania (Microspora), The Journal of Parasitology, Vol. 65, No. 1 (Feb., 1979), pp. 117-122
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Hazard et al. (1979) studied this question.