I. Riddoch, J. Grace, F. E. Fasehun, B. Riddoch, D. O. Ladipo, Photosynthesis and Successional Status of Seedlings in a Tropical Semi-Deciduous Rain Forest in Nigeria, Journal of Ecology, Vol. 79, No. 2 (Jun., 1991), pp. 491-503
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Riddoch et al. (1991) studied this question.
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