Timothy M. Collins, Kenneth Frazer, A. Richard Palmer, Geerat J. Vermeij, Wesley M. Brown, Evolutionary History of Northern Hemisphere Nucella (Gastropoda, Muricidae): Molecular, Morphological, Ecological, and Paleontological Evidence, Evolution, Vol. 50, No. 6 (Dec., 1996), pp. 2287-2304
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Collins et al. (1996) studied this question.