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Pericarp structure has been investigated in five genera of Lamiaceae subg. Nepetoideae tribe Ocimeae which were omitted or poorly studied by Ryding (1992a): Hanceola, Limniboza, Skapanthus, Tetradenia and Thorncroftia . In the light of the results, the systematic position of these and some related genera are discussed. Limniboza is suggested to belong to Ocimeae subtribe Ociminae, the Acrocephalus group. Hanceola (incl. Siphocranion), Isodon, Skapanthus (which is resurrected as a genus), Tetradenia and Thorncroftia apparently do not fit into any of the three traditional subtribes (Hyptidinae, Ociminae or Plectranthinae). The three Asiatic genera, Hanceola, Isodon and Skapanthus might together form a new subtribe. Tetradenia and Thorncroftia have more isolated positions in the Ocimeae.
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Olof Ryding (1993) studied this question.
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