Timothy M. Goater, Allen W. Shostak, Julie A. Williams, Gerald W. Esch, A Mark-Recapture Study of Trematode Parasitism in Overwintered Helisoma anceps (Pulmonata), with Special Reference to Halipegus occidualis (Hemiuridae), The Journal of Parasitology, Vol. 75, No. 4 (Aug., 1989), pp. 553-560
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Goater et al. (1989) studied this question.