Frank S. Gilliam, Bradley M. Yurish, L. M. Goodwin, Community Composition of an Old Growth Longleaf Pine Forest: Relationship of Soil Texture, Bulletin of the Torrey Botanical Club, Vol. 120, No. 3 (Jul. - Sep., 1993), pp. 287-294
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Gilliam et al. (1993) studied this question.
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