"A Comparative Study of Some Mouthpart Adaptations of Uca Annulipes (H. Milne Edwards, 1837) and U. Vocans (Linnaeus, 1758) (Brachyura, Ocypodidae) in Relation to their Habitats" published on 01 Jan 2004 by Brill.
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Shirley S. L. Lim (2004) studied this question.
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