Peter Goldblatt, John C. Manning, Peter Bernhardt, Pollination Biology of Lapeirousia Subgenus Lapeirousia (Iridaceae) in Southern Africa; Floral Divergence and Adaptation for Long-Tongued Fly Pollination, Annals of the Missouri Botanical Garden, Vol. 82, No. 4 (1995), pp. 517-534
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Goldblatt et al. (1995) studied this question.