Johan Elmberg, Petri Nummi, Hannu Poysa, Kjell Sjoberg, Relationships Between Species Number, Lake Size and Resource Diversity in Assemblages of Breeding Waterfowl, Journal of Biogeography, Vol. 21, No. 1 (Jan., 1994), pp. 75-84
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Elmberg et al. (1994) studied this question.
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