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During observational fieldwork in undisturbed ranges of free-roaming bison and moose, I have identified approximately 8% of surface bones as spirally or green-fractured due to documented carnivore activity, and 5% as spirally or green-fractured due to trampling or dust wallowing by bison. The bones of smaller species suffer up to 50% breakage. Bone modifications by wild wolves and bears are briefly described, as are characteristics of fractures caused by trampling and wallowing.
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Gary Haynes (1983) studied this question.
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