Pycreus sect. Tuberculati was created by Chermezon to contain a single species with derived nutlets: P. divulsus, a Malagasy endemic. Kükenthal transferred this species to his new section Muricati. However, a detailed study of the nutlet epidermis shows P. divulsus is not closely related to the other species Kükenthal placed in the latter section. In addition P. divulsus subsp. africanus is upgraded to the species level based on its larger, smooth nutlets and African continental range.
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Reynders et al. (2010) studied this question.
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