The mode of cleavage of the centrolecithal tick eggs has so far been considered to be superficial. Evidence could be obtained from an electron-microscopic study of Omithodoros moubata Murray 1877 that cleavage of this species is total. The early development leads to a diploblastic stage composed of a superficial layer of yolk-free micromeres (ectomesoderm) and a core of yolk-rich macromeres (primary entoderm).
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Fagotto et al. (1988) studied this question.
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