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Permian–Cenozoic Hymenoptera Genomic Radiation, Orussidae Ectoparasitoid Sisterhood to Apocrita, and the 284–0 Ma Total-Evidence Time-Tree of Parasitoidism and Independent Eusociality Across 108 NCBI RefSeq sequence records totaling 1,223,380 bp raw (1,173,240 bp aligned supermatrix: 68 complete mitogenomes, 52 in-frame COX1 coding cores, and 40 nuclear Orco/Vg/Vg-R/Dnmt3/MRJP1/Art3 orthologs + 285 Mkv characters; B = 500 RuNIE replicates), our hybrid RuNIE + *MAFFT-L-INS-i + *IQ-TREE2-LG-C60-F-G4 synthesis resolves the 281.4 Ma radiation of Hymenoptera, confirms a single 226.4 Ma origin of parasitoidism uniting Orussidae + Apocrita, unites ants and bees at 148.6 Ma, and maps four independent Cretaceous origins of eusociality. Canonical Preprint URL: https://crvscience.com/labs/preprints/hymenoptera-radiation-eusociality-timetree-preprint Master (Concept) Zenodo DOI: 10.5281/zenodo.23995669 Open-Source Software Repository (GNU GPL-3.0): https://github.com/bpwhite/crvscience-labs
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Bryan Pinckney White (2026) studied this question.
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