Xenophrys vuquangensis Pham, Le, Nguyen, Pham, Luong, Nguyen, Thai, Ninh, Nguyen & Nguyen, 2026, sp. nov.

Xenophrys vuquangensis sp. nov. (Figs. 4–6) Holotype. VNU A.01 (Field number VQ2025.164), adult male, collected on 12 April 2025 by A. V. Pham in the evergreen forest (18°12'31.5"N, 105°20'07.2"E, at an elevation of 2010 m a.s.l.) within Vu Quang National Park, Ha Tinh Province, Vietnam. Paratypes. VNU A.02 (Field number VQ2025.88), VNU A.03 (Field number VQ2025.146), VNU A.04 (Field number VQ2025.165), VNU A.05 (Field number VQ2025.166), VNU A.06 (Field number VQ2025.167), VNU A.07 (Field number VQ2025.168), and VNU A.08 (Field number VQ2025.177) (Fig 6), adult males, VNU A.09 (Field number VQ2025.141), adult female, the same data as the holotype. Diagnosis. The new species is distinguishable from its congeners by a combination of the following morphological characters: 1) size medium (SVL 63.9–68.6 mm in males; SVL 77.6 mm in the female); 2) tympanum distinct (TD/ ED 0.53–0.56 in males; TD/ED 0.51 in the female); 3) vomerine teeth present; 4) tongue feebly notched posteriorly; 5) external vocal sac present; 6) toes with rudimentary webbing; 7) subarticular tubercles absent; 8) lateral fringes narrow along toes; 9) dorsal skin of body with small granules; 10) flank with numerous large tubercles; 11) dorsum with a dorsal ridge, in>-<shape; 12) belly and ventral surface of thigh with numerous tubercles; 13) dorsal head and back reddish brown with a dark brown triangle between the eyes; 14) a dark brown marking edged, in>-<shape, along dorsal ridge; and 15) flank and back of thigh with larger white spots. Description of holotype. Adult male; SVL 67.6 mm; head wider than long (HL 25.0 mm, HW 26.6 mm); snout round in dorsal view, obtusely protruding in lateral view; nostril lateral, oval, closer to the tip of snout than to the eye (NS 3.9 mm, EN 4.5 mm); canthus rostralis angular; pupil vertically elliptical; loreal region acute, concave; eye diameter shorter than snout length (SL 8.4 mm, ED 7.6 mm); internarial distance narrower than interorbital distance but wider than upper eyelid (IND 6.2 mm, IOD 7.5 mm, UEW 6.0 mm); tympanum distinct, oval, 54% of eye diameter (TD 4.1 mm); vomerine teeth present, maxillary teeth present; tongue round, feebly notched posteriorly; external vocal sac present. Forelimbs: Forelimbs length (HAL 30.2 mm, FLL 16.8 mm); relative finger lengths I<II<IV<III; fingers free of webbing; tips of fingers round, subarticular tubercles and palmar tubercles present. Hindlimbs: Tibia longer than thigh (FeL 31.0 mm, TbL 36.4 mm), approximately 4.5 times longer than wide (TbW 8.1 mm); tips of toes round, relative length of toes I<II<V<III<IV; toes with rudimentary webbing; subarticular tubercles absent; toes with narrow lateral fringes; tarsal fold absent; inner metatarsal tubercle elongate; outer metatarsal tubercle absent; tibio-tarsal articulation reaching to the anterior corner of eye when adpressed along body. Skin texture in life: Dorsal surface with small granules; a small pointed horn-like tubercle present on the outer edge of the upper eyelid; back with a dorsal ridge, in>-<shape; supratympanic fold distinct, curves down broadly through upper tympanum, terminating above forelimbs; dorsolateral folds prominent; flank with numerous large tubercles; dorsal surface of limbs granular with short transverse ridges; throat and chest smooth; belly and ventral surface of thigh with numerous tubercles. Coloration in life: Dorsal head and back reddish brown with a dark brown triangle between the eyes; a dark brown marking edged, in>-<shape, along dorsal ridge; supratympanic fold light cream; dorsolateral folds brown; tympanum dark brown; pupil black, outlined in brown; upper lip with a discontinuous white stripe, running from the nostril to shoulder; flanks yellowish brown with larger white spots; dorsal surface of fore and hind limbs brown with dark crossbars; throat and chest dark brown, pectoral glands white, with two white lines on either side of the jawline; anterior part of belly white with brown pattern, posterior part of belly white yellow with spots; ventral surface of limbs dark brown with white marking, ventral part of thigh, posterior part of thigh reddish brown with white spots; femoral glands white; toe webbing dark brown. Coloration in preservative: Dorsal surface of head and back greyish with a dark brown triangle marking between the eyes; a dark brown marking edged, in>-<shape, along dorsal ridge; upper lip with a discontinuous light stripe; flanks greyish with some larger light spots; throat and chest brown; belly cream with brown pattern; dorsal surface of fore and hind limbs greyish with dark brown crossbars; posterior part of thigh dark brown with larger white spots; femoral gland white; toe webbing dark brown. Variation. Measurements and morphological characters of the type series are given in Table 8. Adult males are smaller than adult females. Dorsum is reddish brown or grayish brown. Distribution. Xenophrys vuquangensis sp. nov. is currently only known from the type locality in Vu Quang National Park, Ha Tinh Province, Vietnam (Fig. 1). Etymology. The specific name “vuquangensis ” is derived from the type locality, Vu Quang National Park in Ha Tinh Province. For the common names we suggest Vu Quang Horned Toad (English) and Cóc mắt vũ quang (Vietnamese). ......continued on the next page Natural history. Specimens were found at night between 19:00 and 22:30, on the ground, on tree branches or on the rocks nearby streams. The surrounding habitat was evergreen forest of medium and large hardwoods mixed with shrubs. Air temperature was 22–28°C and relative humidity was 75–85%. Other amphibian species found at the site were Amolops minhlei Pham, Hoang, Nguyen, Nguyen, Nguyen, Nguyen, Thai, Hoang & Pham, Leptobrachella eos (Ohler, Wollenberg, Grosjean, Hendrix, Vences, Ziegler & Dubois), Leptobrachella sp., Odorrana nasica (Boulenger), Odorrana sp., Kurixalus odontotarsus (Ye & Fei), and Polypedates mutus (Smith).