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Family PTEROBRYACEAE Kindb. Calyptothecium duplicatum (Schwägr.) Broth. (Argent 1973: 574, as C. acutifolium (Brid.) Broth. R). Hildebrandtiella endotrichelloides Müll. Hal. M (Renauld 1898: 296, as H. rotundifolia Renauld R). Hildebrandtiella phleoides (Desv. ex Brid.) B. H. Allen & Magill R (Bridel 1827: 431, as Hypnum phleoides Desv. ex Brid.). Jaegerina formosa Besch. R (Bescherelle 1880a: 263). Jaegerina solitaria (Brid.) A. Jaeger R (Bridel 1812: 90, as Hypnum solitarium Brid.). Jaegerina solitaria var. ramosa Besch. E R (Bescherelle 1880a: 262). Orthostichidium involutifolium subsp. thomeanum (Broth.) Argent (De Sloover 1979: 404). Orthostichidium pentastichum (Brid.) B. H. Allen & Magill M (Palisot de Beauvois 1805: 65, as Hypnum imbricatum P. Beauv. R). Orthostichopsis subimbricata var. borbonica (Renauld & Cardot) Cardot E R (Renauld 1898: 196, as Pilotrichella subimbricata var. borbonica Renauld & Cardot). Orthostichopsis sublivens (Besch.) Broth. E R (Bescherelle 1880a: 268, as Meteorium sublivens Besch.). Pterobryon julaceum Broth. (Ellis et al. 2018: 14).
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Wilding et al. (2026) studied this question.