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Reductodrilus longelytratus Packova & Kundrata sp. nov. Figs 1, 2, 3 Type material. Holotype: male, “ Philippinen Luzon M. Balbalasan // S Boettcher III. 1918 ” (MFNB). Type locality. Philippines: Luzon, Balbalasang [Kalinga province, Balbalasang – Balbalan National Park, Cordillera Central mts., ~ 950 m, 17°29'N, 121°03'E; Bílý and Kubáň 2012]. Diagnosis and comparison. This species can be unambiguously recognized by its relatively large body size (8.7 mm; up to 7.1 mm in all other congeners), antennomeres 4–10 approx. as long as wide (distinctly wider than long in all other congeners; Figs 5 C, 9 D), elytra being 1.7 × as long as wide at humeri (up to 1.45 × in all other congeners) and 3.2 × as long as pronotum (up to 2.5 × in other species; Figs 5 E, 9 E), and by the more or less uniformly lightly colored elytral pubescence (elytral pubescence bicolorous, with major portion of elytra having distinctly darker setae in all other congeners). Description. Male. Body (Fig. 1 A – C) 8.7 mm long from frons to apex of abdomen and 6.6 mm long from frons to apex of elytra, 2.8 mm wide at humeri; yellowish to pale brown, head slightly darker, antennae, palpi, major portions of elytra (except at humeri), major portions of tibiae (except basally), tarsomeres, and sides of abdominal ventrites dark brown; body surface covered with yellowish to pale brown setae. Head (Fig. 2 A) including eyes 0.65 × as wide as pronotum, dorsomedially very slightly convex, generally smooth, sparsely covered with fine punctures and short semi-erect setae mostly oriented anteriorly. Frontoclypeal region frontally abruptly declivous, antennal fossae separated by 0.7 × longitudinal diameter of fossa; rim dorsally forming narrow supra-antennal carina. Eyes moderately large, basally slightly emarginate. Labrum beneath (i. e., not on the same plane with frontoclypeal region), weakly sclerotized, sparsely covered with long suberect to erected setae. Mandibles unidentate, moderately long and robust, sickle-shaped, strongly curved; basally covered with sparse long setae. Maxillary palpi relatively short, 4 - segmented, basal palpomere shorter than wide, palpomere 2 slightly longer than wide; palpomere 3 shortest, wider than long; ultimate palpomere longest, ~ 2 × as long as wide, apically dorsoventrally flattened and obliquely cut. Labial palpi minute, 3 - segmented; palpomere 2 short, transverse; ultimate palpomere longer than wide, similar in shape to ultimate maxillary palpomere. Antenna (Fig. 2 C) with 11 antennomeres, moderately long, almost reaching midlength of elytra (when positioned backwards), rather robust, scape longer than wide, antennomere 2 shortest, approximately as long as wide; antennomeres 3–10 strongly serrate, antennomere 3 ~ 2.2 × as long as antennomere 2, ~ 1.25 × as long as wide, antennomeres 4–10 subequal in shape and length, approx. as long as wide; antennomere 11 slightly longer than antennomere 10, elongate, widest behind midlength, apically rounded. Pronotum (Fig. 2 B) ~ 1.5 × wider than long, slightly convex, widest before posterior angles. Anterior margin slightly rounded, with the edge turned upwards; lateral margins slightly rounded, anteriorly also slightly turned upwards; posterior margin slightly bisinuate, medially slightly produced. Anterior angles inconspicuous; posterior angles very short, almost rectangular, apically narrowly rounded. Surface of disc uneven, with distinct median longitudinal groove, one smaller posteromedial depression, and two shallow anterolateral depressions, moderately densely covered with punctures and setae; punctures moderately large, shallow, intervals between punctures usually 1–2 puncture diameters, interstices rather smooth; setae moderately long, semi-erect, mostly oriented posteriorly, those near posterior margin oriented anteriorly. Lateral pronotal carina complete, visible in dorsal view, anteriorly continuous with anterior margin of pronotum. Hypomeron with surface mostly smooth and shiny, with shallow punctures and semi-erect setae. Prosternum distinctly transverse, with prosternal process reduced to short median protrusion. Scutellar shield (Fig. 2 B) tongue-shaped, large, almost as long as wide, widest anteriorly, gradually slightly narrowed toward apex, apex subtruncate; covered with relatively fine punctures which are approx. the same size as those on elytral bases, and with short semi-erect setae. Mesoventrite short, roughly triangular. Metaventrite with distinct long discrimen; metepisternum distinctly wider basally and gradually narrowed apicad. Elytra (Fig. 1 A, B) elongate, both combined 1.7 × as long as wide, 3.2 × as long as pronotum, dehiscent, not meeting along suture, sides slightly sinuate, apically slightly divergent, apices separately rounded, surface rather uneven and covered with moderately large shallow punctures (smaller on elytral bases) which are subserial to irregularly distributed, and with short, semierect setae which are oriented posteriad. Leg (Fig. 2 D) slender; coxa elongate; femur robust, elongate, flattened, obliquely attached to trochanter, distinctly wider than tibia, with short cavity for reception of tibia; tibia elongate, very slightly longer than femur, apically with two short spurs; tarsus slender, distinctly shorter than tibia in fore legs, less distinctly shorter in mid legs, and only slightly shorter in hind legs; tarsomeres 1–4 progressively shorter, tarsomere 1 longest, longer than combined lengths of tarsomeres 2 and 3, tarsomere 4 shortest, medially emarginate, ventrally with membrane, tarsomere 5 elongate, narrower than preceding tarsomeres; pretarsal claws simple, slightly curved. Abdomen (Figs 1 C, 2 D) with all ventrites densely covered with fine punctures and moderately long, semi-erect, posteriorly directed setae; tergites IX and X (Fig. 3 A) tightly connected by membrane, tergite IX slightly more than twice as wide as long, anteromedially deeply emarginate to receive base of tergite X, densely covered with moderately long setae; tergite X weakly sclerotized, almost as long as wide, apically widely rounded; sternite IX (Fig. 3 B) elongate, ~ 1.8 × as long as wide, gradually slightly widened toward apex, widest subapically, apically subtruncate and medially slightly emarginate. Male genitalia (Fig. 3 C – E) ~ 1.9 × as long as maximum phallobase width; median lobe ~ 2 × as long as phallobase, not surpassing apices of parameres, elongate, slender, subparallel-sided but widest at mid-length, apically rounded, basal struts relatively short and distinctly divergent; parameres elongate, robust, narrowed from 3 / 5 of their length to apex, apically membranous and rounded; phallobase transverse, ~ 1.45 × as wide as long, widely U-shaped. Females and immature stages unknown. Etymology. The species name refers to relatively long elytra of this species compared to its congeners. Distribution. Philippines.
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