Poa pseudobulbosa Bor

3.Poa pseudobulbosaBor, Notes Roy. Bot. Gard. Edinburgh 31 (3): 395. 1972. = Poa diversifolia var. crassipes Hack. ex Boiss., Fl. Orient. 5: 601. 1884. Type Protologue Libano ad Cedros., Bl. 3889. Type specimen. Libanos aux Cedros, 7 Jun 1855, Blanche 3889. (lectotype, here designated: (G [G 00785172]; isolectotype: W-Hackel [W 1916-0011101, image!], JE [JE 00005087, image!]). Type. Turkey • C 4, Antalya, NW side of Ak Dağ (S of Geyik Dağ), by the little lake, 2,200–2,300 m, shady rocks; [28 Aug 1947], Davis 14342 a. (lectotype, here designated [with Bor’s original description]: K [K 000496556]); isolectotypes: ANK, K [K 000496557]. Note. The stated holotype does not appear to exist at E. Description. Perennials; hermaphroditic; tufted, without rhizomes or stolons, tufts dense, slender, moderate sized, green to pale green; tillers intravaginal and extravaginal; prophylls retrorsely scabrous; bulbs present, slightly inflated at the base. Culms (16 –) 20–43 cm, erect, slender, several per tuft, smooth or lightly scabrous below the panicle, smooth or moderately densely retrorse scaberulous below the nodes, glabrous. Leaves mostly basal; leaf sheaths strongly imbricated at base, smooth or scaberulous, glabrous; basal sheaths forming bulbs, the outer ones slightly inflated, loose, thin papery, watery-white to eventually brown and withering to form a weak but distinct tunic, the inner ones somewhat thickened (bulb slender, elongate, gradually tapered upward); flag - leaf sheaths 5–9 cm long, margins fused 50–65 % the length; ligules 0.5–0.9 (– 1.3) mm long, hyaline, or watery-white, abaxially scabrous, apices truncate and erose, sparsely asperous; ligules of sterile shoots 0.3–0.6 mm long; blades 0.5–4 cm long, and 1–1.5 mm broad, 0.5–0.8 mm diam.; culm blades very short, less than 1 / 2 their sheath in length, folded with involute margins, moderately firm, abaxially smooth, adaxially sparsely scabrous along veins, margins sparsely scabrous, apex narrowly prow-tipped, scabrous, abaxially smooth; flag - leaf blade erect, the collar indistinct; blades of sterile shoots 2–7 cm long. Panicles (3 –) 4–8.5 cm long, 1–2.5 cm wide, loosely contracted, very sparsely flowered (9–20 spikelets); branches (1 –) 2 at the lowest nodes; primary branches 0.8–3.1 cm, 0.17–0.38 x the panicle, av. 0.3 x with 1–3 few spikelets per branch, ascending, weakly angled sparsely to moderately scabrous; pedicels mostly slightly shorter than their spikelets, Spikelets 5–7 mm long, narrowly lanceolate, not bulbiferous; florets 1–3; rachilla internodes 1.2–1.7 mm long, visible in lateral view, glabrous, densely muriculate; glumes unequal, keels scabrous distally, margins narrowly hyaline, edges scaberulous distally, apices narrowly acute, sharp pointed; lower glumes 3–3.5 mm long, lanceolate, 3 - veined; upper glumes 3.5–4 mm long (broadly elliptic when flattened), 3–5 - veined, margins and apex hyaline; callus glabrous; lemmas 3–4.7 mm long, 5 - veined, narrowly lanceolate, violaceous, distinctly keeled, glabrous, keel scabrid, surfaces faintly muriculate to sparsely scaberulous, intermediate veins faint to moderately distinct extending to near the apex, upper margins narrowly hyaline, apices acute, sharply pointed or slightly erose, infrequently mucronulate; palea shorter than the lemma, keels distally scabrous, intercostal zone muriculate. Flowers bisexual; anthers 1.9 mm long. Caryopses 2.5 mm long, distinctly sulcate, hard, glabrous, tightly adherent to the palea and lemma; hilum 0.4 mm long, round to oval. Clade code: Hph. Distribution and ecology. Lebanon, Syria, Turkey. Upper mountain to low alpine mesic meadows, on calcareous substrates. Rarely collected; in Turkey, 2,180–2,300 m. Conservation status. Listed as Near Threatened (IUCN 2013). Local. Uncommon. Fairly rare but local populations seemingly stable. Comments. Compared to Poa anatolica and P. girdevensis, P. pseudobulbosa is taller and narrower panicled (20–43 cm tall, 1–2.5 cm wide), with branches 0.17–0.38 x the panicle length (vs. 0.44–0.78 x the panicle length) and lower glumes longer, 3–3.5 mm (vs. 2–3 mm). Representative specimens. Turkey. Antalya (Davis 14342 a ANK, E, K; 14557 E; 16557 E); (Soreng et al. 8863). Isparta (Soreng et al. 4139; 4141 - a *; 8246 * ISTE 33941, US). Lebanon. (Type; haut Lebanon supra Entre Diman et Yamouny, Blanche 3889 - bis G). Syria. (Dschebel-eck-Scheik, Bertschinger 1937 G). Names of Uncertain Application to Poa ser. Bulbosae, etc., and a new record for Turkey of Poa supina