M. Debussche, J. Cortez, I. Rimbault, Variation in Fleshy Fruit Composition in the Mediterranean Region: The Importance of Ripening Season, Life-Form, Fruit Type and Geographical Distribution, Oikos, Vol. 49, No. 3 (Jul., 1987), pp. 244-252
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Debussche et al. (1987) studied this question.
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