M. Proulx, F. R. Pick, A. Mazumder, P. B. Hamilton, D. R. S. Lean, Experimental Evidence for Interactive Impacts of Human Activities on Lake Algal Species Richness, Oikos, Vol. 76, No. 1 (May, 1996), pp. 191-195
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Proulx et al. (1996) studied this question.