S. L. Pimm, R. L. Kitching, The Determinants of Food Chain Lengths, Oikos, Vol. 50, No. 3, Trophic Exploitation, Community Structure and Dynamics of Cyclic and Non-Cyclic Populations. Proceedings of a Symposium Held 25-27 May, 1985, at Hällnäs, Sweden (Nov., 1987), pp. 302-307
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Pimm et al. (1987) studied this question.
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