Michael E. Jacoby, Grant V. Hilderbrand, Christopher Servheen, Charles C. Schwartz, Stephen M. Arthur, Thomas A. Hanley, Charles T. Robbins, Robert Michener, Trophic Relations of Brown and Black Bears in Several Western North American Ecosystems, The Journal of Wildlife Management, Vol. 63, No. 3 (Jul., 1999), pp. 921-929
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Jacoby et al. (1999) studied this question.