Charnov and Schaffer's (1973) model for annual versus perennial reproduction in plants has been extended to include both density dependence and environmental variability. Two models of density dependence were investigated, and the method of invasibility analysis (Turelli 1978) was used to study their behavior. The results are interpreted in terms of bet-hedging theory; it is shown that the criticisms of this interpretation by Hastings and Caswell (1979) are unfounded.
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Michael Bulmer (1985) studied this question.
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