Jerrold I. Davis, Paul S. Manos, Isozyme Variation and Species Delimitation in the Puccinellia nuttalliana Complex (Poaceae): An Application of the Phylogenetic Species Concept, Systematic Botany, Vol. 16, No. 3 (Jul. - Sep., 1991), pp. 431-445
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Davis et al. (1991) studied this question.
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