Answers are grounded in this paper and cite their sources.
Melville H. Hatch; The Natural Classification of the Families of Coleoptera, Annals of the Entomological Society of America, Volume 49, Issue 1, 1 January
No takes yet. Share an insight, caveat, or question.
Melville H. Hatch (1956) studied this question.
Synapse has enriched 10 related papers. Consider them for comparative context: