Morphology and Phylogeny of the Reduvioidea (Hemiptera: Heteroptera). Part II. Wing Venation1

Evidences of the identity of the veins of the cubitalanal field of the hemipterous wing are given. In the fore wing of auchenorrhynchous Homoptera these veins are: Cm (anterior to the clavus) ; Cu2 (closely associated with the claval furrow) ; Pcu, 1st A, and 2d A (in the clavus). In Heteroptera Cu2 is absent, and the clavus contains only Pcu and 1st A. There is no anal lobe in either suborder, the clavus representing the postcubital region of the fore wing. In the hind wing of Auchenorrhyncha the veins of the cubital-anal field are the same as in the fore wing; a cubital furrow lies just behind Cu2, and the anal fold lies just behind 1st A. In Heteroptera this region of the hind wing is similar, except that Cu2 is absent, and the cubital furrow often is bifid and may contain secondary veins between its branches. In both Auchenorrhyncha and Heteroptera the anal lobe of the hind wing usually is well developed, and the jugal lohe is always absent. In the Cimicomorpha the hind-wing venation of the Miridae, Nabidae, Microphysidae, Anthocoridae, Joppeicidae, Thaumastocoridae, and Velocipedidae is of the same basic type, characterized particularly by the absence of Sc and by having a Y- or V-shaped pattern of Pcu and 1st A. The tingids resemble this group, but have a distinct Sc. The nabids and microphysids seem exceptional in sometimes having R and M separated distally, as in the Pentatomomorpha. The reduviid venation is distinct from that of the other Cimicomorpha, and is best characterized by a narrow postcubital sector containing subparallel Pcu and 1st A veins. Venation is useful in distinguishing subfamilies of the Nabidae. Anthocoridae, and Reduviidae; in the Reduviidae it serves in characterizing three major groups of subfamilies.