After 100 years of burial in moist well aerated sand, 21 seeds of Verbascum blattaria, one seed of Verbascum thapsus and one seed of Malva rotundifolia, germinated and produced normal plants. Six additional seeds, of the original 50 seeds of each of 20 varieties, germinated and died before identification was possible.
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Kivilaan et al. (1981) studied this question.
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