Sandra Van Oevelen, Els Prinsen, Rupert De Wachter, Elmar Robbrecht, The Taxonomic Value of Bacterial Symbiont Identification in African Psychotria (Rubiaceae), Systematics and Geography of Plants, Vol. 71, No. 2, Plant Systematics and Phytogeography for the Understanding of African Biodiversity (2001), pp. 557-563
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Oevelen et al. (2001) studied this question.
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