Dave I. Thompson, Trevor J. Edwards, Johannes van Staden, In Vitro Germination of Several South African Summer Rainfall Disa (Orchidaceae) Species: Is Seed Testa Structure a Function of Habitat and a Determinant of Germinability?, Systematics and Geography of Plants, Vol. 71, No. 2, Plant Systematics and Phytogeography for the Understanding of African Biodiversity (2001), pp. 597-606
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Thompson et al. (2001) studied this question.
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