Answers are grounded in this paper and cite their sources.
H. K. Buechner, I. O. Buss, W. M. Longhurst, A. C. Brooks, Numbers and Migration of Elephants in Murchison Falls National Park, Uganda, The Journal of Wildlife Management, Vol. 27, No. 1 (Jan., 1963), pp. 36-53
No takes yet. Share an insight, caveat, or question.
Buechner et al. (1963) studied this question.
Synapse has enriched 10 related papers. Consider them for comparative context: