Thomas Alerstam, Gudmundur A. Gudmundsson, Kerstin Johannesson, Resources for Long Distance Migration: Intertidal Exploitation of Littorina and Mytilus by Knots Calidris Canutus in Iceland, Oikos, Vol. 65, No. 2 (Nov., 1992), pp. 179-189
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Alerstam et al. (1992) studied this question.
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