Nigel P. Barker, Peter H. Weston, John P. Rourke, Gail Reeves, The Relationships of the Southern African Proteaceae as Elucidated by Internal Transcribed Spacer (ITS) DNA Sequence Data, Kew Bulletin, Vol. 57, No. 4 (2002), pp. 867-883
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Barker et al. (2002) studied this question.
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