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Glenn D. DelGiudice, L. David Mech, Ulysses S. Seal, Patrick D. Karns, Winter Fasting and Refeeding Effects on Urine Characteristics in White-Tailed Deer, The Journal of Wildlife Management, Vol. 51, No. 4 (Oct., 1987), pp. 860-864
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DelGiudice et al. (1987) studied this question.
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