A further chromosomally distinct sibling species of the Helophorus minutus complex from Morocco, with additional notes on Spanish H. calpensis Angus, 1988 (Coleoptera: Helophoridae)

Helophorus atlantis sp. n. is described from the Moyen Atlas region of Morocco (Ifrane and Fes). Morphologically it appears indistinguishable from the more lightly built species of the H. minutus group (H. minutus Fabricius, 1775 and H. calpensis Angus, 1988 Angus, R. B. 1988. ‘A new sibling species of Helophorus F. (Coleoptera, Hydrophilidae) revealed by chromosome analysis and hybridisation experiments’. Aquatic Insects, 10: 171–183. [Taylor & Francis Online], [Web of Science ®] , [Google Scholar]), and its karyotype is most similar to that of H. calpensis, but with important differences, especially in the sex chromosomes. An additional karyotype of H. calpensis, from the Coto Doñana, Provincia de Huelva, Spain provides a major extension to the known distribution of this species, hitherto known only from its type locality (Tarifa, Provincia de Cádiz) and also confirms the chromosomal distinction between H. calpensis and H. atlantis.