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Dale R. McCullough, Floyd W. Weckerly, Pamela I. Garcia, Rand R. Evett, Sources of Inaccuracy in Black-Tailed Deer Herd Composition Counts, The Journal of Wildlife Management, Vol. 58, No. 2 (Apr., 1994), pp. 319-329
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McCullough et al. (1994) studied this question.
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