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A female entered a swarm of males of Anarete pritchardi and coupled sequentially with two of the males. The horizontal and vertical movements of the midges prior to, during, and following the couplings were recorded on high speed movies and analyzed. The couplings lasted 3.3 and 6.4 sec. Upon disengaging, the male took off before the female did.
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Chiang et al. (1978) studied this question.
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