Experimental analysis shows root development varies by soil type and solution concentration.
Ponderosa pine occurs naturally on a wide variety of soils and consequently the establishment of reproduction varies greatly. The author grew seedlings in eight soil types varying from light pumice to clays, and found that they grew best on lightly combined soils. In a brief note supplementing this article the author reports his work on the influence of soil solution concentration on root development. The more diluted the solution the more extensive the root system must be.
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James O. Howell (1932) studied this question.
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