Douglas S. Cram, Ronald E. Masters, Fred S. Guthery, David M. Engle, Warren G. Montague, Northern Bobwhite Population and Habitat Response to Pine-Grassland Restoration, The Journal of Wildlife Management, Vol. 66, No. 4 (Oct., 2002), pp. 1031-1039
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Cram et al. (2002) studied this question.